Cranial anatomy of the aetosaur Paratypothorax andressorum Long & Ballew, 1985, from the Upper Triassic of Germany and its bearing on aetosaur phylogeny

Abstract

Riojasuchus tenuisceps Bonaparte 1967 is currently known from four specimens, includingtwo complete skulls, collected in the late 1960s from the upper levels of the Los ColoradosFormation (Late Triassic), La Rioja, Argentina. Computed tomography (CT) scans of theskulls of the holotype and a referred specimen of Riojasuchus tenuisceps and the repreparationof the latter allows recognition of new features for a detailed analysis of its cranialanatomy and its comparison with a wide variety of other archosauriform taxa. The diagnosisof Riojasuchus tenuisceps is emended and two autapomorphies are identified on the skull:(1) a deep antorbital fossa with its anterior and ventral edges almost coinciding with thesame edges of the maxilla itself and (2) a suborbital fenestra equal in size to the palatinepterygoidfenestra. Also, the first digital 3D reconstruction of the encephalon of Riojasuchustenuisceps was carried out to study its neuroanatomy, showing a shape and cranial nervedisposition consistent to that of other pseudosuchians.