New vertebrates from the Ituzaingó Formation (Late Miocene of Entre Ríos Province, Argentina), including first records of Leptodactylus (Amphibia, Anura) and Chelonoidis (Testudines, Cryptodira)

Abstract

The Late Miocene vertebrate fauna from the Entre Ríos Province, Argentina, comes from two units: the marine Paraná Formation and the Lower Member of Ituzaingó Formation (LMIF). This latter includes a lower conglomerate set, with fine quartz gravel, and clay and chalcedony clasts, and fossil remains, traditionally known as ‘Mesopotamiense’ or ‘Conglomerado osífero’. We present new fossil vertebrate remains from the LMIF at the Puerto Víboras locality (Hernandarias, Entre Ríos Province) including Leptodactylus sp. (Amphibia, Anura), Crocodylia indet. Caimaninae indet. and Paranasuchus gasparinae (Alligatoridae), Chelidae indet. (Testudines, Pleurodira), Chelonoidis sp. (Testudines, Cryptodira), Plohophorus paranensis (Cingulata, Glyptodontidae), and Macraucheniinae indet. (Macraucheniidae, Litopterna). The record of Leptodactylus sp. represents the oldest record of the genus and the first amphibian record for the Neogene of the Entre Ríos Province. The material described herein and assigned to Chelonoidis sp. allows to confirm for the first time the presence of a testudinid in the Late Miocene of the Entre Ríos Province.